Mix the warmed buttermilk and melted butter together in a large liquid measuring cup. Combine 4 cups of flour, sugar, yeast and salt together in a standing mixer fitted with dough hook (or you can use a large bowl and mix with a wooden spoon or electric handheld mixer).

With the mixer on low speed, add the buttermilk mixture and eggs and mix until the dough comes together, about 2 minutes. Increase the mixer to medium speed and knead the dough until it is smooth and elastic, about 10 minutes (knead for 15-18 minutes by hand). If after 5 minutes of kneading, the dough is still overly sticky, add up to 1/4 cup more flour, 1 tablespoon at a time until the dough clears the sides of the bowl but has a slight tacky feel when pressed between your fingertips.

Place the dough in a large, lightly greased bowl and cover the top tightly with plastic wrap. Let the dough rise in a warm place until doubled, around 2 to 2 ½ hours.

Combine all the filling ingredients (reserving 1/2 teaspoon orange zest for the frosting) in a small bowl and mix well. Set aside.

When the dough is ready, turn it out onto a lightly floured counter and press it into about a 16 by 12-inch rectangle. Gently brush the filling mixture over the rectangle, using the back of a spoon or rubber spatula.

Lift the longest edge closest to you and begin rolling the dough into a tight log. Pinch the seam closed and roll the log so it is seam side down. Using floss, cut the log into 12 evenly sized rolls. Arrange the rolls on a lined or lightly greased baking pan. Cover the rolls with lightly greased plastic wrap. Let the rolls rise in a warm place until doubled, 1 to 1 ½ hours.

Bake the rolls at 350 degrees for 22-25 minutes, until the rolls are lightly golden on top and cooked through.

Beat together the frosting ingredients in a large bowl. (Start with 1 tablespoon of fresh orange juice and add a little more at a time until desired consistency is reached.) Once rolls are slightly cool spread on top of rolls.

Store in an airtight container.